He couldnt be careful enough on that upstroke.
他扬起翅膀时，真是够小心的了。
decreased upstroke velocity of action potential
动作电位上升速度减慢
The thrust force is mainly due to the drag of the wings in the upstroke.
平衡身体阻力的推力主要由翅膀上拍中产生的阻力提供。
During the course upstroke, the membrane potential attains zero, and an electrical gradient no longer exists.
在升支发展过程中，膜电位一达到零，电位梯度即不再存在。
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